This municipality is located at the centre of the valley. Its name derives from the Roman road whose layout nowadays follows its main street. It has very well preserved remains and a Roman bridge. The Market Square or the Severiano Masides Street are particularly interesting.

La Garganta is located in the foothills of the Sierra de Candelario. It is the highest urban area of the Ambroz Valley at an altitude over 1,000 metres above sea level, where the presence of snow is common in winter. It preserves an old bell for snow. The tradition of livestock farming has linked this village to the Iberian wolf, and it is worth a visit to its creative interpretation centre and to a trap for hunting wolves.

Hervás is the business activity centre. Its well preserved Jewish Quarter was declared a Historic-Artistic Set in 1996 and it belongs to the Network of Jewish Quarters in Spain. The mount Castañar Gallego is also a Protected Landscape. Hervás offers the visitors a great gastronomic and leisure variety, as well as shops with traditional products.

The stone Celtic Bull dominates the village main square and it is the municipality’s symbol. Segura preserves remains from the Vettones, a templar castle and beautifull streets with ashlar stone houses, sloppy narrow streets with charming corners and façades covered with colourful flowers. The Castaños del Temblar are a unique group of giant trees. The town also has a beautiful natural pool in summer.